All these photos are made by digital camera Nikon Coolpix 4500. I am very thankful to prof. Medvedev (curator in Zoological Institute of Russian Academy of Sciences, St.-Petersburg) for opprtunity to use some specimens and to Angrey Yakovlev for technical support.
Made by Konstantin A. Grebennikov

Staphylinidae of Lower Volga Region:

Page 1 (Omaliinae): Anthophagus caraboides, Arpedium quadratum, Lesteva punctata, Olophrum assimile, Omalium caesum, Xylodromus concinnus.

Page 2 (Oxytelinae 1): Anotylus insecatus, Anotylus mendus, Anotylus rugosus, Anotylus sculpturatus, Aploderus caelatus, Bledius debilis, Bledius debilis, Bledius fracticornis, Bledius furcatus, Bledius spectabilis, Bledius tricornis, Bledius unicornis.

Page 3 (Oxytelinae 2): Carpelimus sp., Deleaster dichrous, Elonium pennifer, Manda mandibularis, Oxytelus migrator, Oxytelus sculptus, Platystethus cornutus, Platystethus nodifrons, Platystethus spinosus.

Page 4 (Steninae): Stenus bimaculatus, Stenus boops, Stenus clavicornis, Stenus comma, Stenus fornicatus, Stenus juno, Stenus ludyi, Stenus palposus.

Page 5 (Paederinae 1): Achenium depressum, Achenium quadriceps , Astenus filiformis, Astenus pulchellus, Astenus immaculatus, Cryptobium egregium, Lathrobium boreale, Lathrobium elongatum, Lathrobium flavipes, Lathrobium fovulum, Lathrobium punctatum, Lathrobium rufonitidum, Lathrobium scutellare, Lathrobium taxi.

Page 6 (Paederinae 2): Lithocharis nigriceps, Luzea cephalica, Medon fusculus, Medon mersinus, Paederus fuscipes, Paederus litoralis, Paederus riparius, Pseudomedon obsoletus.

Page 7 (Paederinae 3): Rugilus orbiculatus, Rugilus rufipes, Rugilus similis, Scimbalium anale, Scopaeus bicolor, Scopaeus debilis, Scopaeus gladifer, Scopeaus laevigatus, Sunius fallax, Sunius sp., Throbalium kochi, Leptobium gracile.

Page 8 (Xantholininae): Gyrohypnus fracticornis, Leptacinus batychrus, Leptacinus sulcifrons, Leptacinus linearis, Leptolinus versicolor, Nudobius lentus, Xantholinus distans, Xantholinus roubali, Platyprosopus elongatus, Othius punctulatus.

Page 9 (Staphylininae 1): Erichsonius cinerascens, Gabrius osseticus, Gabrius suffragani, Neobisnius procerulus, Philonthus binotatus, Philonthus cruentatus, Philonthus dimidiatipennis, Philonthus dimidiatus, Philonthus discoideus, Philonthus ebeninus.

Page 10 (Staphylininae 2): Philonthus ephippius, Philonthus fumarius, Philonthus micans , Philonthus rectangulus, Philonthus salinus, Philonthus spinipes, Philonthus succicola, Philonthus tenuicornis, Philonthus velatipennis, Philonthus wuestoffi, Rabigus formosus, Jurechekia asphaltina.

Page 11 (Staphylininae 3): Creophilus maxillosus, Ocypus nitens, Ocypus picipennis, Staphylinus caesareus, Tasgius falcifer, Tasgius inderiensis, Platydracus stercorarius, Acylophorus glaberrimus, Quedius limbatus, Quedius puncticollis, Quedius umbrinus, Heterothops minutus, Heterothops tanygnathoides, Velleius dilatatus.

Page 12 (Tachyporinae): Bolitobius castaneus, Carphacis striatus, Lordithon lunulatus, Leucoparyphus silphoides, Sepedophilus obtusus, Sepedophilus testaceus, Tachyporus chrysomelinus, Tachyporus nitidulus, Tachinus discoideus.

Page 13 (Aleocharinae, Scaphidiinae): Myllaena intermedia, Oligota parva, Homalota plana, Gygronoma dimidiata, Gyrophaena joyi, Atemeles emarginatus, Falagria sulcata, Falagria thoracica, Drusilla canaliculata, Mocyta fungi, Oxypoda abdominalis, Chilopora rubicunda, Aleochara sareptana, Aleochara (Baryodma) sp., Scaphidium quadrimaculatum, Scaphisoma agaricinum.

Staphylinidae of Russia and adjacent countries:

Page 1 (Omaliinae): Cylletron nivale - Buryatiya, Eucnecosum brachypterum - E Sayany, Geodromicus sp. - Kabardino-Balkariya, Geodromicus sp. - NW Caucasus, Lesteva cordicollis - E Sayany, Lesteva sp. - Uzbekistan, Mannerheimia divergens - Buryatiya. Pycnoglypta baicalica - Irkutsk, Xylodromus sp. - Tuva, Xylodromus sp. (undescribed) - Abkhaziya.

Page 2 (Oxytelinae, Piestinae, Trigonurinae, Apatecinae, Olisthaerinae, Oxyporinae): Euphanias pliginskii (Oxytelinae) - Krym, Siagonum humerale (Piestinae) - NW Caucasus, Trigonurus asiaticus (Trigonurinae) - NW Caucasus, Nodynus leucofasciatus (Apatecinae) - Kunashir, Olisthaerus substriatus (Olisthaerinae) - Leningrad Reg., Oxyporus mannerheimi (Oxyporinae) - Kalinin, Oxyporus maxillosus - Kalinin, Oxyporus rufus (Oxyporinae) - Estonia.

Page 3 (Steninae, Xantholininae, Tachyporinae, Aleocharinae, Pselaphinae): Stenus pflegeri (Steninae) - NW Caucasus, Stenus stigmula NW - Caucasus, Stenus velleris - NW Caucasus, Allolinus peliopterus (Xantholininae) - Buryatiya, Tachinus signatus (Tachyporinae) - Bashkiriya, Bolitochara bella (Aleocharinae) - St.-Petersburg, Leptusa sp. (Aleocharinae) - NW Caucasus, Ocalea sp. (Aleocharinae) - NW Caucasus, Batrisotes sp. (Pselaphinae) - NW Caucasus.

Page 4 (Paederinae): Domene sp. - Russian Far East, Lobrathium reitteri - NW Caucasus, Homaeotarsus 2 spp. - Azerbiajan, Lathrobium brunnipes - St.-Petersburg, Lathrobium cavicola - Slovenia, Lathrobium eppelsheimi - NW Caucasus, Lathrobium lomnickii - NW Caucasus, Lathrobium longulum - Rostov Reg., Lathrobium pallidipenne - Kabardino-Balkariya, Nazeris pallidipes - NW Caucasus, Paederus ruficollis - N Caucasus.

Page 5 (Staphylininae 1): Cafius xantholoma - Krym, Philonthus cyanipennis - NW Caucasus, Philonthus decorus - St.-Petersburg, Philonthus marginatus - Bashkiriya, Philonthus psyllophagus - Irkutsk Reg., Remus filum - Krasnodar Terr., Abemus chloropterus - Moldova, Dinothenarus arrosus - NW Caucasus, Dinothenarus flavocephalus - N Caucasus, Dinothenarus sibiricus - Kirghizia, Phucobius simulator - Primor'e.

Page 6 (Staphylininae 2): Emus hirtus - W Europe, Hadropinus fossor - Primor'e, Ocypus cerceticus - NW Caucasus, Ocypus curtipennis - Azerbaijan, Ocypus nitens - Krasnodar Terr., Ocypus sp. - Kirghizia, Ocypus sp. - Japan, Ocypus ophthalmicus - Belgorod Reg.

Page 7 (Staphylininae 3): Ontholestes tesselatus - St.-Petersburg, Ontholestes haroldi - Moldova, Ontholestes murinus - Novgorod, Ontholestes sp. - Primor'e, Platydracus dauricus - Buryatiya, Platydracus fulvipes - St.-Petersburg, Tasgius depressus - NW Caucasus, Tasgius pedator - NW Caucasus , Tasgius sp. - Kunashir, Tasgius sp. (undescribed) - Kirghizia, Astrapaeus ulmi - N Caucasus, Quedius fuliginosus - Kabardino-Balkariya.

Staphylinidae of the world: Borolinus sp. (Osoriinae) - India, Osoriinae spp. (3) - Vietnam, Steninae sp. - Vietnam, "Sterculia sp." (Xantholininae) - Peru, Belonuchus arrogans (Staphylininae) - Columbia, ? Belonuchus sp. - Peru, Platydracus sp. (Staphylininae) - Angola, Platydracus sp. - "British East Africa", Platydracus sp. - Ethiopia, Platydracus sp. - China, Yun-Nan, Creophilus sp. (Staphylininae) - Australia, Creophilus sp. (Staphylininae) - Brazil, Creophilus sp.(Staphylininae) - New Zealand.

Carabidae (of Lower Volga Region mostly):

Page 1: Omophron limbatum, Cicindela silvatica, Cicindela germanica, Cicindela lunulata nemoralis, Cicindela campestris, Cicindela chiloleuca, Cicindela contorta, Cicindela elegans, Cicindela atrata, Cicindela lacteola.

Page 2: Cicindela soluta, Cicindela sahlbergi, Carabus stscheglovi, Carabus estreicheri, Carabus granulatus, Carabus clathratus, Carabus cancellatus, Carabus marginatus, Carabus nemoralis, Carabus hortensis.

Page 3: Carabus glabratus, Carabus hungaricus, Carabus bessarabicus, Carabus convexus, Carabus sexcarinatus, Carabus prometeus, Carabus caucasicus, Cychrus caraboides, Leistus ferrugineus.

Page 4: Notiophilus palustris, Notiophilus biguttatus, Elaphrus uliginosus, Elaphrus cupreus, Lorocera pilicornis, Scarites bucida, Clivina ypsilon, Dischirius chalceus, Asaphidion pallipes.

Page 5: Bembidion argenteolum, Bembidion dalmatinum, Bembidion dentellum, Bembidion glabrum, Bembidion lampros, Bembidion ruficorne, Bembidion striatum, Bembidion tetracolum, Bembidion ustum, Bembidion varium, Bembidion velox.

Page 6: Trechus secalis, Trechus quadristriatus, Pogonus cumanus, Pogonistes angustus, Badister unipustulatus, Badister meridionalis, Licinus cassideus, Callistus lunatus, Cardioderus chloroticus.

Page 7: Chlaenius aeneocephalus, Chlaenius cruralis, Chlaenius spoliatus, Chlaenius vestitus, Chlaenius tristis, Chlaenius coeruleus, Poecilus lepidus, Pterostichus crenuliger, Pterostichus puncticollis, Pterostichus nitens.

Page 8: Pterostichus cursor, Pterostichus oblongopunctatus, Pterostichus inquinatus, Pterostichus longicollis, Agonum viridicupreum, Agonum longiventre, Agonum dorsale, Amara ovata, Amara famelica, Amara apricaria, Amara brunnea, Curtonotus convexiusculus, Curtonotus aulicus.

Page 9: Daptus vittatus, Stenolophus discophorus, Stenolophus scrimshiranus, Anthracus consputus, Dichirotrichus sp., Lebia menetriesi, Lebia humeralis, Demetrias imperialis, Dromius quadraticollis, Drypta dentata.

Page 10: Cymindis lineata, Cymindis variolosa, Polystichus connexus, Brachynus sclopeta, Brachynus bipustulatus, Brachynus hamatus, Mastax thermarum.

Scarabaeidae (of Lower Volga Region):

Page 1: Geotrupes stercorarius, Aphodius distinctus, Aphodius fimetarius, Aphodius ictericus, Aphodius immundus, Aphodius lugens, Aphodius serotinus, Aphodius sturmi.

Page 2: Gymnopleurus mopsus, Onthophagus gibbulus, Onthophagus furcatus, Caccobius schreberi, Oniticellus fulvus, Amphicoma bombyliformis, Oryctes nasicornis, Pentodon bidens, Pentodon idiota.

Page 3: Adoretus nigrifrons, Anomala dubia, Anomala kirgisica, Anomala errans, Blitopertha lineolata, Anisoplia austriaca, Anisoplia segetum, Anisoplia agricola, Anisoplia zwicki, Anisoplia deserticola.

Page 4: Melolontha hippocastani, Polyphylla fullo, Polyphylla alba, Anoxia pilosa, Lasiopsis caninus, Rhizotrogus aestivus, Miltotrogus vernus, Miltotrogus aequinactialis , Chioneosoma astrachanicum, Chioneosoma pulvereum, Amphimallon solsticialis.

Page 5: Maladera holosericea, Homaloplia spireae, Hoplia parvula, Valgus hemipterus, Epicometis hirta, Oxythyrea funesta, Cetonia aurata, Cetonia metallica, Cetonia cuprina, Cetonia hungarica, Cetonia karelini.